This report concerns vascular plants observed and collected during the 1998 Ralph Brown/Royal Geographical Society Expedition to the Volhynian Polissia (NW Ukraine). The author participated in the first stage of the Expedition in July 1998, Team 2. The route was Kovel' - Buzaky - Island near Khoteshiv - Bank of the Prypiat River near Shchytyn - Bank of the Sviate lake near Zalukhiv - Bank of the Bile lake near Nevir - Kovel' (5--18 July 1998). Of course, the term of the study was not sufficient for a complete floristic survey of the area. However, basic data on the flora of vascular plants were collected.
The main source of information for the list of species was Expedition records of the author. Data of other members of the Expedition team were used when possible. In addition to that, for some species herbarium (KW) and literature information was involved. If we consider the number of species of vascular plants occurring in the region, Polissya can be regarded as the most floristically depauperate natural zone of Ukraine. Of course, it cannot be compared to such floristically rich areas as the Carpathians, Crimea, or even to some still preserved natural areas in the forest-steppe and steppe zones of Ukraine.
However, the natural vegetation of Polissya is among the best preserved in Ukraine. Because of that many Polissian areas are very interesting from the floristical viewpoint. In particular, the Western Polissya is interesting for botanists because of its weird mixture of floristic (phytogeographic) elements, including western (Western and Central European), northern (Boreal, Arctoboreal, etc.), montane (Carpathian, pan-Alpine, Boreoalpine, Arcto-Alpine) ones.
The list of species encountered during the Expedition is provided below. It should not be regarded as the complete list of the vascular flora of the region, since many important areas were left unexplored during the 1998 Expedition. However, it is a rather good "sample" of a typical Polissian flora. Data on interesting and noteworthy floristic finds are also provided in reports of other members of the Expedition, in publications being prepared, and in available literature.
The list is divided into four major parts: Ferns and fern allies (Lycopodiophyta, Equisetophyta, Polypodiophyta), gymnosperms (Pinophyta), and angiosperms (monocots and dicots separately). Almost all other taxa are listed alphabetically (families within classes; genera within families, and species within genera). Species not observed during the Expedition, but which are definitely present in the region, are marked with asterisk (*). Some species are marked with "?", which means that the species has not been observed by the author, but probably occur in the region.
Lycopodiaceae
Lycopodium annotinum L.
Lycopodium clavatum L.
Isoetaceae
Isoetes lacustris L. [Collected by Vakarenko and Dubyna]
Equisetaceae
Equisetum arvense L.
Equisetum fluviatile L. [= E. heleocharis Ehrh.]
*Equisetum hyemale L.
Equisetum palustre L.
*Equisetum pratense L.
*Equisetum sylvaticum L.
Athyriaceae
Athyrium filix-femina (L.) Roth
Cystopteris fragilis (L.) Bernh.
Dennstaedtiaceae [= Hypolepidaceae]
Pteridium aquilinum (L.) Kuhn
Dryopteridaceae [= Aspidiaceae p.p.]
Dryopteris carthusiana (Vill.) H.P. Fuchs
Dryopteris cristata (L.) A. Gray
Dryopteris filix-mas (L.) Schott
Ophioglossaceae [incl. Botrychiaceae]
Ophioglossum vulgatum L.
Thelypteridaceae
Thelypteris palustris Schott
Pinaceae
Picea abies (L.) Karst. [Widely cultivated as forestry plantations; exclaves of the native spruce also occur in the region]
Pinus sylvestris L.
Cupressaceae
Juniperus communis L.
Alismataceae
*Alisma lanceolatum With.
Alisma plantago-aquatica L.
*Caldesia parnassifolia (L.) Parl. [Can be expected in the region]
Sagittaria sagittifolia L.
Alliaceae
Allium angulosum L.
Allium oleraceum L.
Allium podolicum (Aschers. & Graebn.) Blocki ex Racib. [ ~ Allium paniculatum s.l.]
Allium scorodoprasum L.
Amaryllidaceae
*?Galanthus nivalis L. [A spring plant. Not observed in July]
Araceae
Acorus calamus L.
Calla palustris L.
Butomaceae
Butomus umbellatus L.
Cyperaceae
Bolboschoenus maritimus (L.) Palla
Carex acuta L.
*Carex acutiformis Ehrh.
Carex appropinquata [Reported by Vakarenko]
Carex cinerea Poll. [Reported by Vakarenko]
Carex contiqua Hoppe
Carex digitata L.
Carex echinata Murr.
Carex elata All. [s.l., incl. subsp. omskiana (Meinsh.) Jalas]
*Carex ericetorum Poll. [A common plant of the Polissian pine forests. However, no Expedition records are available and this species was probably overlooked]
Carex flava L. [s.l., incl. C. flavella V. Krecz.]
Carex hirta L.
Carex juncella (Fries) Th. Fries [s.l., incl. Carex wiluica Meinsh.]
Carex lasiocarpa Ehrh. [Reported by Vakarenko]
Carex muricata L.
Carex nigra (L.) Reichard
Carex otrubae Podp.
Carex ovalis L. [Carex leporina L., p.p., nom. ambig.]
Carex panicea L. [Reported by Vakarenko]
Carex praecox Schreb.
Carex pseudocyperus L.
Carex remota L.
Carex riparia Curt. [Reported by Vakarenko]
Carex rostrata Stokes
*Carex tomentosa L.
Carex vesicaria L.
Carex vulpina L.
Cyperus fuscus L.
Eleocharis acicularis (L.) Roem. & Schult.
Eleocharis palustris (L.) Roem. & Schult.
Eleocharis uniglumis (Link) Schult.
Eriophorum polystachyon L.
Eriophorum vaginatum L.
Rhynchospora alba (L.) Vahl [Collected by Vakarenko in August 1999]
Scirpus lacustris L. [= Schoenoplectus lacustris (L.) Palla]
*Scirpus radicans Schkuhr
Scirpus sylvaticus L.
Hydrocharitaceae
Elodea canadensis Michx.
Hydrocharis morsus-ranae L.
Stratiotes aloides L.
Iridaceae
Iris pseudacorus L.
Juncaceae
Juncus alpino-articulatus Chaix [Juncus alpinus Vill. nom. illeg.]
Juncus articulatus L.
Juncus bufonius L. [s.l., incl. Juncus ambiguus Guss., Juncus ranarius Song. & Perrier ex Billot]
*Juncus bulbosus L. [Most probably occurs in the area]
Juncus compressus Jacq.
Juncus conglomeratus L.
Juncus effusus L.
Juncus gerardii Loisel.
Juncus inflexus L.
Juncus squarrosus L. [Collected by Vakarenko]
Juncus tenuis Willd.
Luzula multiflora (Ehrh.) Lej.
Luzula pilosa (L.) Willd.
Juncaginaceae
Triglochin palustre L. [Reported by Vakarenko]
Lemnaceae
Lemna minor L.
Lemna trisulca L.
Spirodela polyrrhiza (L.) Schleid.
Liliaceae [s.l., incl. Trilliaceae, Asparagaceae, etc.]
Anthericum ramosum L.
Asparagus officinalis L.
Convallaria majalis L.
*Gagea lutea (L.) Ker-Gawl. [A spring plant]
*Gagea minima (L.) Ker-Gawl. [A spring plant]
Lilium martagon L.
Maianthemum bifolium (L.) F.W. Schmidt
Paris quadrifolia L.
Polygonatum multiflorum (L.) All.
Polygonatum odoratum (Mill.) Druce
*Scilla bifolia L. [A spring plant]
Veratrum album L. [s.l., incl. subsp. lobelianum (Bernh.) Schuebl. & Martens = Veratrum lobelianum Bernh.]
Najadaceae
Najas marina L. [s.l.]
Orchidaceae
?Dactylorhiza fuchsii (Druce) Soo [Plants in fruits, identification tentative]
Dactylorhiza incarnata (L.) Soo
Epipactis helleborine (L.) Crantz
Neottia nidus-avis (L.) Rich.
Platanthera bifolia (L.) Rich.
Poaceae
Agrostis canina L.
Agrostis gigantea Roth
Agrostis stolonifera L.
Agrostis tenuis Sibth.
Alopecurus aequalis Sobol.
Alopecurus geniculatus L.
Alopecurus pratensis L.
Anisantha tectorum (L.) Nevski
Apera spica-venti (L.) P. Beauv.
Arrhenatherum elatius (L.) J. Presl & C. Presl
Avena sativa L.
Avenella flexuosa (L.) Drejer
Beckmannia eruciformis (L.) Host
Brachypodium pinnatum (L.) P. Beauv.
Brachypodium sylvaticum (Huds.) P. Beauv.
Briza media L.
Bromopsis inermis (Leyss.) Holub
Bromus mollis L.
Bromus squarrosus L.
Calamagrostis arundinacea (L.) Roth
Calamagrostis canescens (Web.) Roth
Calamagrostis epigeios (L.) Roth
Catabrosa aquatica (L.) P. Beauv.
Corynephorus canescens (L.) P. Beauv.
Cynosurus cristatus L.
Dactylis glomerata L.
Deschampsia cespitosa (L.) P. Beauv.
Digitaria ischaemum (Schreb.) Muehl.
Digitaria sanguinalis (L.) Scop.
Echinochloa crus-galli (L.) P. Beauv.
Elytrigia repens (L.) Nevski
Eragrostis minor Host
Eragrostis pilosa (L.) P. Beauv.
Festuca altissima All.
Festuca arundinacea Schreb.
Festuca beckeri (Hack.) Trautv. [incl. Festuca beckeri (Hack.) Trautv. subsp. polesica (Zapal.) Tzvel.]
Festuca gigantea (L.) Vill.
Festuca ovina L. [sensu latissimo]
Festuca pratensis Huds.
Festuca rubra L. [s.l.]
Glyceria fluitans (L.) R. Br.
Glyceria maxima (C. Hartm.) Holmberg [Glyceria aquatica (L.) Wahlenb.]
Glyceria notata Chevall. [Glyceria plicata (Fries) Fries]
Hierochloë odorata (L.) P. Beauv.
Holcus mollis L.
Koeleria cristata (L.) Pers. [s.l.]
Koeleria glauca (Spreng.) DC. [s.l.]
Koeleria grandis Bess. ex Gorski
Leersia oryzoides (L.) Sw.
Lolium perenne L.
Melica nutans L.
Milium effusum L.
Molinia caerulea (L.) Moench
Nardus stricta L.
Phalaroides arundinacea (L.) Rausch.
Phleum phleoides (L.) Karst.
Phleum pratense L.
Phragmites australis (Cav.) Trin. ex Steud. [Phragmites communis Trin.]
Poa angustifolia L.
Poa annua L.
*Poa bulbosa L.
Poa compressa L.
Poa nemoralis L.
Poa palustris L.
Poa pratensis L.
Poa trivialis L.
Poa turfosa Litv. [Poa palustris L. subsp. volhynensis (Klok.) Tzvel. s. str.]
Setaria glauca (L.) P. Beauv.
Setaria viridis (L.) P. Beauv.
Sieglingia decumbens (L.) Bernh.
Potamogetonaceae
Potamogeton acutifolius Link
Potamogeton berchtoldii Fieb.
Potamogeton compressus L.
Potamogeton crispus L.
Potamogeton lucens L.
Potamogeton natans L.
Potamogeton nodosus Poir.
*Potamogeton obtusifolius Mert. & Koch
Potamogeton pectinatus L.
Potamogeton perfoliatus L.
?Potamogeton pusillus L.
Scheuchzeriaceae
Scheuchzeria palustris L. [Reported by Vakarenko and Heluta]
Sparganiaceae
Sparganium emersum Rehm.
Sparganium erectum L. [s.l., incl. subsp. microcarpum (Newm.) Domin and subsp. neglectum (Beeby) K. Richt.]
Typhaceae
Typha angustifolia L.
Typha latifolia L.
Aceraceae
Acer campestre L.
*Acer negundo L.
Acer platanoides L.
Acer pseudoplatanus L.
Acer tataricum L.
Adoxaceae
Adoxa moschatellina L.
Amaranthaceae
Amaranthus albus L.
Amaranthus retroflexus L.
Apiaceae [Umbelliferae]
Aegopodium podagraria L.
Aethusa cynapium L.
Angelica archangelica L. [= Archangelica officinalis]
Angelica palustris (Bess.) Hoffm. [= Ostericum palustre]
Angelica sylvestris L.
Anthriscus sylvestris (L.) Hoffm.
Archangelica officinalis Hoffm.
Berula erecta (Huds.) Coville [= Siella erecta]
Calestania palustris (L.) K.-Pol.
Cenolophium denudatum (Hornem.) Tutin [= Cnidium fischeri Spreng.]
Chaerophyllum temulum L.
Cicuta virosa L.
Cnidium dubium (Schkuhr) Thell.
Daucus carota L.
Eryngium planum L.
Heracleum sibiricum L.
Heracleum sphondylium L.
Hydrocotyle vulgaris L. [Reported by Heluta & Vakarenko]
Laserpitium prutenicum L.
Oenanthe aquatica (L.) Poir.
Ostericum palustre (Bess.) Bess.
Pimpinella saxifraga L.
Sanicula europaea L.
Siella erecta (Huds.) M. Pimen.
Sium latifolium L.
Torilis japonica (Houtt.) DC.
Xanthoselinum alsaticum (L.) Schur [= Peucedanum alsaticum L.]
Apocynaceae
*Vinca major L. [Cultivated and escaped]
Aristolochiaceae
Aristolochia clematitis L.
Asarum europaeum L.
Asclepiadaceae
Vincetoxicum hirundinaria Medik.
Asteraceae
Achillea collina J. Becker ex Reichenb.
Achillea millefolium L. [s.l.]
*Antennaria dioica (L.) Gaertn.
*Anthemis arvensis L.
Arctium lappa L.
Arctium minus (Hill) Bernh.
Arctium tomentosum Mill.
Artemisia absinthium L.
Artemisia campestris L.
Artemisia vulgaris L.
Bidens cernua L.
Bidens frondosa L.
Bidens tripartita L.
Carduus acanthoides L. [s.l.]
Carduus crispus L.
Centaurea cyanus L.
Centaurea diffusa Lam. [Kovel', railroad]
Centaurea jacea L.
Centaurea rhenana Boreau [incl. Centaurea rhenana Boreau subsp. pseudomaculosa (Dobrocz.) Dostal = Centaurea pseudomaculosa Dobrocz.]
Chondrilla juncea L.
Cichorium intybus L.
Cirsium arvense (L.) Scop.
Cirsium oleraceum (L.) Scop.
Cirsium palustre (L.) Scop.
Cirsium rivulare (Jacq.) All.
Cirsium vulgare (Savi) Ten.
Conyza canadensis (L.) Cronq. [Erigeron canadensis L.]
Crepis biennis L.
Crepis tectorum L.
Cyclachaena xanthiifolia (Nutt.) Fresen.
Erigeron acris L. [s.l.]
Eupatorium cannabinum L.
Filago arvensis L.
Galinsoga parviflora Cav.
Gnaphalium sylvaticum L.
Gnaphalium uliginosum L.
Helichrysum arenarium (L.) Moench
Hypochoeris maculata L.
Hypochoeris radicata L.
Inula britannica L.
Inula germanica L.
Inula hirta L.
Inula salicina L.
*Jurinea cyanoides (L.) Reichenb.
Lactuca quercina L. [s.l., incl. Lactuca quercina L. subsp. chaixii (Vill.) Celak.]
Lactuca serriola L.
Lapsana communis L.
Leontodon autumnalis L.
Leontodon danubialis Jacq.
Leontodon hispidus L.
Lepidotheca suaveolens (Pursh) Nutt.
Leucanthemum vulgare Lam. [s.l., incl. Leucanthemum vulgare Lam. subsp. ircutianum (Turcz. ex DC.) Tzvel. = Leucanthemum ircutianum Turcz. ex DC.]
Matricaria recutita L.
Mycelis muralis (L.) Dumort.
*Onopordum acanthium L.
Petasites hybridus (L.) Gaertn., Mey. & Scherb.
Phalacroloma annuum (L.) Dumort. [s.l.]
Picris hieracioides L.
Ptarmica cartilaginea (Ledeb. ex Reichenb.) Ledeb.
Ptarmica salicifolia (Bess.) Serg.
*Pyrethrum corymbosum (L.) Scop. [Probably occurs in the area]
Senecio erucifolius L.
Senecio vulgaris L.
Solidago virgaurea L.
Sonchus arvensis L.
Sonchus oleraceus L.
Tanacetum vulgare L.
Taraxacum officinale Wigg. [aggr.]
Tragopogon orientalis L.
Tripleurospermum inodorum (L.) Sch. Bip.
Tussilago farfara L.
*Xanthium albinum (Widd.) H. Scholz [Probably occurs as a weed]
Balsaminaceae
Impatiens noli-tangere L.
Impatiens parviflora DC.
Berberidaceae
*Berberis vulgaris L. [Cultivated in settlements]
Betulaceae
Betula pendula Roth
Betula pubescens Ehrh. (Betula verrucosa Ehrh.)
Boraginaceae
Asperugo procumbens L.
Buglossoides arvensis (L.) Johnst.
Cynoglossum officinale L.
Echium vulgare L.
Myosotis arvensis (L.) Hill
Myosotis nemorosa Bess.
Myosotis palustris (L.) L.
Myosotis micrantha Pall. ex Lehm.
Myosotis sylvatica Ehrh. ex Hoffm.
Pulmonaria angustifolia L.
Pulmonaria obscura Dumort. [Pulmonaria officinalis L. aggregate]
Symphytum officinale L.
Brassicaceae [Cruciferae]
Alliaria petiolata (M. Bieb.) Cavara & Grande
Arabidopsis thaliana (L.) Heynh.
Arabis planisiliqua (Pers.) Reichenb.
*Armoracia rusticana Gaertn., Mey. & Scherb. [Cultivated and escaped]
Barbarea stricta Andrz.
Barbarea vulgaris R. Br.
Berteroa incana (L.) DC.
Brassica campestris L.
Capsella bursa-pastoris (L.) Medik.
Cardamine amara L.
Cardamine dentata Schult. [= Cardamine palustris (Wimm. & Graebn.) Peterm.]
Cardamine impatiens L.
Cardamine parviflora L.
Cardaminopsis arenosa (L.) Hayek
Cardaria draba (L.) Desv. [Rare, along railroad in Kovel']
Dentaria bulbifera L.
Descurainia sophia (L.) Webb ex Prantl
Draba nemorosa L.
Erophila verna (L.) Bess.
Erysimum cheiranthoides L.
Lepidium densiflorum Schrad.
Lepidium ruderale L.
Raphanus raphanistrum L.
Rorippa amphibia (L.) Bess.
Rorippa austriaca (Crantz) Bess.
Rorippa palustris (L.) Bess.
Rorippa sylvestris (L.) Bess.
Sisymbrium altissimum L. [Rare weed in Kovel']
Sisymbrium loeselii L.
Sisymbrium officinale (L.) Scop.
Thlaspi arvense L.
Turritis glabra L.
Callitrichaceae
Callitriche sp. [No fruits available]
Campanulaceae
Campanula bononiensis L.
Campanula cervicaria L.
Campanula glomerata L. [s.l., incl. Campanula farinosa Andrz.]
Campanula patula L.
Campanula persicifolia L.
Campanula rotundifolia L.
Campanula trachelium L.
Jasione montana L.
Cannabaceae
Cannabis sativa L. [incl. Cannabis ruderalis Janisch.]
Humulus lupulus L.
Caprifoliaceae
Sambucus nigra L.
Sambucus racemosa L.
*Viburnum opulus L.
Caryophyllaceae
Arenaria serpyllifolia L. [s.l., incl. A. uralensis Pall. ex Spreng.]
Cerastium holosteoides Fries
Cerastium semidecandrum L. [s.l., incl. Cerastium rotundatum Schur]
Coccyganthe flos-cuculi (L.) Fourr. [= Coronaria flos-cuculi (L.) A. Br. = Lychnis flos-cuculi L.]
Cucubalus baccifer L.
Dianthus borbasii Vandas
Dianthus deltoides L.
Dianthus pseudosquarrosus (Novak) Klok. [D. arenarius L. aggr.]
Dianthus stenocalyx Juz. [D. superbus L. aggr.]
Eremogone saxatilis (L.) Ikonn.
Gypsophila fastigiata L.
Herniaria glabra L.
Herniaria polygama J. Gay
Holosteum umbellatum L.
Melandrium album (Mill.) Garcke
Melandrium dioicum (L.) Coss. & Germ.
Myosoton aquaticum (L.) Moench
Oberna behen (L.) Ikonn. [= Silene vulgaris (Moench) Garcke]
Otites borysthenicus (Grun.) Klok.
Psammophiliella muralis (L.) Ikonn.
Sagina nodosa (L.) Fenzl
Sagina procumbens L.
Saponaria officinalis L.
Scleranthus annuus L.
Scleranthus perennis L.
Silene chlorantha (Willd.) Ehrh.
Silene lithuanica Zapal.
Silene nutans L.
Spergula arvensis L.
Spergularia rubra (L.) J. Presl & C. Presl
Stellaria alsine Grimm. [= Stellaria uliginosa Murr.]
Stellaria holostea L.
Stellaria graminea L.
Stellaria media (L.) Vill.
Stellaria palustris Retz.
Steris viscaria (L.) Rafin. [= Viscaria viscosa (L.) Achers. = Viscaria vulgaris Bernh.]
Celastraceae
Euonymus europaea L.
Euonymus verrucosa Scop.
Ceratophyllaceae
Ceratophyllum demersum L.
Ceratophyllum submersum L.
Chenopodiaceae
Atriplex patula L.
Atriplex sagittata Borkh. [Atriplex nitens Schkuhr, Kovel', railroad]
Atriplex tatarica L.
Chenopodium album L.
Chenopodium glaucum L.
Chenopodium opulifolium Schrad. ex DC. [Rare, Kovel']
Chenopodium rubrum L.
Chenopodium strictum Roth
Kochia scoparia (L.) Schrad.
Salsola tragus L. [s.str., Salsola ruthenica Iljin, nom. illegit., Salsola australis R. Br., p.p.]
Convolvulaceae
Calystegia sepium (L.) R. Br.
Convolvulus arvensis L.
Cornaceae
Swida sp. [Cultivated, Kovel']
Corylaceae
Corylus avellana L.
Crassulaceae
Sedum acre L.
Sedum ruprechtii (Jalas) Omelcz. [Hylotelephium ruprechtii (Jalas) Tzvel.]
Cuscutaceae
Cuscuta europaea L. [s.l.]
Cuscuta lupuliformis Krock.
Dipsacaceae
Knautia arvensis (L.) Coult.
Scabiosa ochroleuca L.
Droseraceae
*Aldrovanda vesiculosa L. [Probably occurs in the area]
Drosera anglica Huds.
Drosera rotundifolia L.
Elatinaceae
Elatine sp. [No fruits available]
Ericaceae [see also Vacciniaceae]
Andromeda polifolia L.
*Arctostaphylos uva-ursi (L.) Spreng. [Reported by Vakarenko and Heluta]
Calluna vulgaris (L.) Hull
Ledum palustre L.
Euphorbiaceae
Euphorbia cyparissias L.
Euphorbia esula L. [s.l.]
?Euphorbia palustris L.
Euphorbia virgata Waldst. & Kit. [incl. Euphorbia virgultosa Klok.]
?Euphorbia volhynica Bess. ex M. Raciborsky
Mercurialis perennis L.
Fabaceae [Leguminosae]
Anthyllis macrocephala Wend.
Chamaecytisus austriacus (L.) Link
Chamaecytisus ruthenicus (Fisch. ex Woloszcz.) Klaskova
Coronilla varia L.
Genista tinctoria L.
Lathyrus niger (L.) Bernh.
Lathyrus pratensis L.
Lathyrus sylvestris L.
Lathyrus vernus (L.) Bernh.
Lotus corniculatus L. [aggr., incl. Lotus arvensis Pers.]
Medicago falcata L. [aggr.]
Medicago lupulina L.
Medicago sativa L. [Cultivated and escaped]
Melilotus albus Medik.
Melilotus officinalis (L.) Pall.
Trifolium alpestre L.
Trifolium arvense L.
Trifolium aureum Poll.
Trifolium campestre Schreb.
Trifolium dubium Sibth.
Trifolium hybridum L.
Trifolium medium L.
Trifolium montanum L.
Trifolium prantense L.
Trifolium repens L.
Vicia angustifolia Reichard
Vicia dumetorum L.
Vicia sepium L.
Vicia tetrasperma (L.) Schreb.
Fagaceae
Quercus robur L.
Fumariaceae
*Corydalis bulbosa (L.) DC. [Corydalis solida (L.) Clairv., a spring plant]
*Corydalis cava (L.) Schweigg. & Koerte [A spring plant]
Gentianaceae
Centaurium erythraea Rafn
*Centaurium uliginosum (Waldst. & Kit.) G. Beck ex Ronn.
*Gentiana pneumonanthe L.
Geraniaceae
Erodium cicutarium (L.) L'Her.
Geranium palustre L.
Geranium pratense L.
Geranium robertianum L.
Geranium sanguineum L.
Geranium sylvaticum L.
Grossulariaceae
Grossularia reclinata (L.) Mill.
Ribes nigrum L.
Haloragaceae
Myriophyllum spicatum L.
Hippuridaceae
Hippuris vulgaris L.
Hypericaceae
Hypericum montanum L.
Hypericum perforatum L.
Lamiaceae [Labiatae]
Acinos arvensis (Lam.) Dandy
Ajuga genevensis L.
Ajuga reptans L.
Ballota nigra L. [s.l.]
Betonica officinalis L.
Clinopodium vulgare L.
Galeobdolon luteum Huds.
Galeopsis bifida Boenn.
Galeopsis pubescens Bess.
Galeopsis tetrahit L.
Glechoma hederacea L.
Lamium album L.
Lamium maculatum (L.) L.
Lamium purpureum L.
Leonurus quinquelobatus Gilib. ex Usteri
Lycopus europaeus L.
Lycopus exaltatus L. fil.
Mentha aquatica L.
Mentha arvensis L.
Mentha longifolia (L.) Huds.
Mentha x verticillata L. [= Mentha aquatica x Mentha arvensis]
Origanum vulgare L.
Prunella vulgaris L.
Salvia pratensis L.
Scutellaria galericulata L.
Scutellaria hastifolia L.
Stachys palustris L.
Stachys sylvatica L.
Teucrium chamaedrys L.
Teucrium scordium L.
Thymus x oblongifolius Opiz [= Thymus x polessicus Klok. = Thymus pulegioides x Thymus serpyllum]
Thymus pulegioides L. [aggr., incl. Thymus ovatus Mill.]
Thymus serpyllum L.
Lentibulariaceae
Pinguicula vulgaris L.
Utricularia vulgaris L.
Linaceae
Linum catharticum L. [Cathartolinum catharticum (L.) Small]
Loranthaceae
Viscum album L.
Lythraceae
Lythrum salicaria L.
Lythrum virgatum L.
Peplis portula L.
Malvaceae
Lavatera thuringiaca L.
Malva excisa Reichenb.
Malva neglecta Wallr.
Malva pusilla Smith
Malva sylvestris L.
Menyanthaceae
Menyanthes trifoliata L.
Nymphaeaceae
Nuphar lutea (L.) Smith
Nymphaea alba L.
Nymphaea candida J. Presl & C. Presl
Oleaceae
*Fraxinus excelsior L. [Cultivated]
*Ligustrum vulgare L. [Cultivated]
Onagraceae
Chamerion angustifolium (L.) Holub
Epilobium hirsutum L.
Epilobium montanum L.
Epilobium palustre L.
Oenothera biennis L. [s.l.]
Orobanchaceae
Orobanche sp.
Oxalidaceae
Oxalis acetosella L.
Xanthoxalis stricta (L.) Small [Xanthoxalis fontana (Bunge) Holub]
Papaveraceae
Chelidonium majus L.
*Papaver rhoeas L.
Plantaginaceae
Plantago arenaria Waldst. & Kit.
Plantago lanceolata L.
Plantago major L.
Plantago media L.
Polemoniaceae
?Polemonium caeruleum L.
Polygalaceae
Polygala amarella Crantz
Polygonaceae
Bistorta major S.F. Gray [Polygonum bistorta L.]
Fallopia convolvulus (L.) A. Love [Polygonum convolvulus L.]
Fallopia dumetorum (L.) Holub [Polygonum dumetorum L.]
Persicaria amphibia (L.) S.F. Gray [Polygonum amphibium L.]
Persicaria hydropiper (L.) Spach [Polygonum hydropiper L.]
Persicaria lapathifolia (L.) S.F. Gray [s.l., Polygonum lapathifolium L.]
Persicaria minor (Huds.) Opiz [Polygonum minus Huds.]
Polygonum aviculare L. [s.l.]
?Rumex acetosa L. [Records need confirmation. Probably R. thyrsiflorus]
Rumex acetosella L.
?Rumex aquaticus L. [No fruits available, identification tentative]
Rumex confertus Willd.
Rumex crispus L.
Rumex hydrolapathum Huds.
Rumex maritimus L.
Rumex obtusifolius L.
Rumex thyrsiflorus Fingerh. [Rumex acetosa L. subsp. thyrsiflorus (Fingerh.) Hayek]
Primulaceae
Anagallis arvensis L.
Hottonia palustris L.
Lysimachia nummularia L.
Lysimachia vulgaris L.
Naumburgia thyrsiflora (L.) Reichenb.
Trientalis europaea L.
Pyrolaceae
Orthilia secunda (L.) House
Pyrola chlorantha Swartz
Pyrola rotundifolia L.
Ranunculaceae
Actaea spicata L.
Anemone sp. [Spring plants. The following three species most probably occur in the area: Anemone nemorosa L., Anemone ranunculoides L., Anemone sylvestris L.]
Batrachium aquatile (L.) Dumort.
Batrachium trichophyllum (Chaix) Bosch
Caltha palustris L.
Clematis recta L.
Ficaria verna Huds.
?Hepatica nobilis Mill. [A spring species. Probably occurs in the area]
Pulsatilla sp. [Spring species]
Pulsatilla patens (L.) Mill. [incl. P. latifolia Rupr.]
Pulsatilla pratensis (L.) Mill. [incl. P. nigricans Stoerck]
Ranunculus acris L.
Ranunculus flammula L.
Ranunculus lingua L.
Ranunculus polyanthemos L.
Ranunculus repens L.
Ranunculus sceleratus L.
Thalictrum flavum L.
Thalictrum lucidum L.
Thalictrum minus L.
Rhamnaceae Juss.
Frangula alnus Mill.
?Rhamnus cathartica L.
Rosaceae
Agrimonia eupatoria L.
Alchemilla sp.
Comarum palustre L.
Crataegus curvisepala Lindm. [s.l., incl. Crataegus pseudokyrtostyla Klok.]
Crataegus fallacina Klok.
Filipendula denudata (J. Presl & C. Presl) Fritsch [= Filipendula ulmaria (L.) Maxim. subsp. denudata (J. Presl & C. Presl) Hayek]
Filipendula ulmaria (L.) Maxim.
Filipendula vulgaris Moench [Filipendula hexapetala Gilib.]
Fragaria vesca L.
Fragaria viridis Duch.
Geum rivale L.
Geum urbanum L.
Malus sylvestris Mill.
Padus avium Mill.
Potentilla alba L.
Potentilla anserina L.
Potentilla argentea L.
Potentilla erecta (L.) Raeusch.
Potentilla impolita Wahlenb.
Pyrus communis L.
Rosa canina L.
Rosa majalis Herrm.
Rubus caesius L.
Rubus idaeus L.
Rubus nessensis W. Hall
Sanquisorba officinalis L.
Sorbus aucuparia L.
Rubiaceae
Cruciata glabra (L.) Ehrend.
Cruciata laevipes Opiz
Galium aparine L.
Galium boreale L. [s.l.]
Galium maximum G. Moris
Galium mollugo L.
Galium odoratum (L.) Scop.
Galium palustre L.
Galium rivale (Sibth. & Smith) Griseb.
Galium verum L.
Salicaceae
Populus alba L.
Populus nigra L.
Populus tremula L.
Salix acutifolia Willd.
Salix alba L.
Salix aurita L.
Salix caprea L.
Salix cinerea L.
Salix fragilis L.
Salix myrtilloides L.
Salix pentandra L.
Salix rosmarinifolia L.
Salix triandra L.
Saxifragaceae
Chrysosplenium alternifolium L.
Scrophulariaceae
?Digitalis grandiflora Mill. [Probably occurs in the area]
Euphrasia brevipila Burn. & Gremli
Euphrasia parviflora Schag.
Euphrasia stricta D. Wolff ex J.F. Lehm.
Gratiola officinalis L.
Limosella aquatica L.
Linaria vulgaris Mill.
Melampyrum arvense L.
Melampyrum cristatum L.
Melampyrum nemorosum L. [s.l., incl. Melampyrum polonicum (Beauverd) Soo]
Melampyrum pratense L.
Odontites vulgaris Moench [= Odontites serotina (Lam.) Dum.]
Rhinanthus aestivalis (N. Zing.) Schischk. & Serg.
Rhinanthus minor L.
Scrophularia nodosa L.
Scrophularia umbrosa Dumort.
Verbascum lychnitis L.
Verbascum nigrum L.
Verbascum phlomoides L.
Verbascum thapsus L.
Veronica anagallis-aquatica L.
Veronica anagalloides Guss.
Veronica beccabunga L.
Veronica chamaedrys L.
Veronica dillenii Crantz
Veronica incana L.
Veronica longifolia L.
Veronica officinalis L.
Veronica scutellata L.
Veronica spicata L. [s.l.]
Veronica teucrium L.
Solanaceae
Lycium barbarum L. [Cultivated (and escaped?) in Kovel']
Solanum dulcamara L.
Solanum nigrum L.
Thymelaeaceae
?Daphne mezereum L. [Probably occurs in the area. Needs confirmation]
Tiliaceae
Tilia cordata Mill.
Trapaceae
?Trapa natans L. [sensu lato, no definite records available]
Ulmaceae
Ulmus glabra Huds.
Urticaceae
Urtica dioica L.
Urtica galeopsifolia Wierzb. ex Opiz
Urtica urens L.
Vacciniaceae
Oxycoccus palustris Pers.
Vaccinium myrtillus L.
Vaccinium uliginosum L.
Vaccinium vitis-idaea L. [Rhodococcum vitis-idaea (L.) Avror.]
Valerianaceae
Valeriana officinalis L. [s.l.]
Violaceae
Viola arvensis Murr.
*Viola hirta L. [A spring plant]
Viola mirabilis L.
*Viola odorata L. [A spring plant]
?Viola reichenbachiana Jord. ex Boreau
?Viola riviniana Reichenb.
Viola tricolor L. [s.l., incl. Viola matutina Klok.]
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